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Dicranopteris linearis var. tomentosa

(Luerss.) Miles K.Thomas (2025)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Gleicheniales Family: Gleicheniaceae Genus: Dicranopteris

hairy uluhe, staghorn fern, uluhe, unuhe

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Bibliography

Name Published In: American Fern J., 115(1):89-90 (2025)

Other References

Palmer 2003:115 (K, O, Mo, L, M, H [as D. linearis f. emarginata]); Thomas & Gallaher 2023:213 (COMBNOV&STATNOV, Syn. D. linearis f. emarginata = D. linearis var. emarginata); Thomas 2025:89 (COMBNOV, Syn. D. linearis var. emarginata = D. linearis var. tomentosa)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Dicranopteris linearis var. tomentosa National Park, Makaopuhi Crater Preserved_Specimen Stone, B.C. 2982 Hawaii BISH 1959-06-26
2 Dicranopteris linearis var. tomentosa unknown Preserved_Specimen (Not on sheet) 3 BISH
3 Dicranopteris linearis var. tomentosa unknown Preserved_Specimen (Not on sheet) BISH
4 Dicranopteris linearis var. tomentosa S. Kona, Alika Homesteads, koa mill road Metrosideros forest Preserved_Specimen St.John, H. 18541 Hawaii BISH 1937-12-27
5 Dicranopteris linearis var. tomentosa Kaala. Northern Waianae Mountains. Ridgeline near HECO tower near the summit. Growing in cloud forest dominated by Metrosideros polymorpha, Hydrangea arguta, Cheirodendron platyphyllum, Dryopteris glabra, Bazzania, Plagiochila, Pyrrhobryum, Leucobryum, Asplenium contiguum, Lycopodium venustulum, Peperomia macraeana, Rubus rosifolius, Adenophorus tamariscinus, Begonia foliosa, and others. Preserved_Specimen Thomas, M.K. 591 Oahu BISH 2023-07-25
6 Dicranopteris linearis var. tomentosa Kilauea with Metrosideros, Lycopodium cernuum, Cibotium, Sadleria Preserved_Specimen Degener, O. 9179 Hawaii BISH 1922-12-15
7 Dicranopteris linearis var. tomentosa Mt. Kalena, Waianae Metrosideros forest Preserved_Specimen Grant, M.L. 7385 Oahu BISH 1934-09-30
8 Dicranopteris linearis var. tomentosa Kehena Ditch trail, near bottom 1st gulch W of Honkaenui with Metrosideros, Cheirodendron, Pelea, Vaccinium, Spaghnum, Dubautia, Gouldia, Straussia, Clermontia, Gunnera, Cyanea Preserved_Specimen Gillett, G.W. 1903 Hawaii BISH 1966-06-26
9 Dicranopteris linearis var. tomentosa above Hiilawe woods Preserved_Specimen Rock, J.F.C. 4586 Hawaii BISH 1909-07-18
10 Dicranopteris linearis var. tomentosa upper Waiakea FR, 1/2 mi E of tree planting road boggy; with Metrosideros collina, Cibotium glaucum, Dicranopteris linearis, Juncus Preserved_Specimen Higashino, P.K. 7411 Hawaii BISH 1978-05-09
11 Dicranopteris linearis var. tomentosa Puna, Kikala-Keokea Homesteads open Metrosideros forest Preserved_Specimen St.John, H. 22277 Hawaii BISH 1946-12-22
12 Dicranopteris linearis var. tomentosa down Chain of Craters Road ~ 9 mi toward Kalapana from intersection with Crater Rim Rd; up old Kalapana Trail ~0.3 mi., 10 m to L of trail with Metrosideros, Dicranopteris Preserved_Specimen Newell, C.L. 562 Hawaii BISH 1966-08-23
13 Dicranopteris linearis var. tomentosa Kilauea scrub Preserved_Specimen Degener, O. 9178 Hawaii BISH 1929-01-01
14 Dicranopteris linearis var. tomentosa above Hiilawe woods Preserved_Specimen Rock, J.F.C. 4583 Hawaii BISH 1909-07-18
15 Dicranopteris linearis var. tomentosa Kaala, Piko Trail sunny bank Preserved_Specimen Topping, D.L. 3755 Oahu BISH 1934-05-29
16 Dicranopteris linearis var. tomentosa North Kohala District, Kalala ahupuaa, Lahikiola cinder cone In understory of Metrosideros woodland Preserved_Specimen Bunch, P.M. 105 Hawaii BISH 1989-03-13
17 Dicranopteris linearis var. tomentosa summit of Mt. Kaala woods Preserved_Specimen Grether, D.F. 3922 Oahu BISH 1945-10-31
18 Dicranopteris linearis var. tomentosa Kaala, summit swamp Preserved_Specimen Takeuchi, W.N. 2347 Oahu BISH 1985-07-21
19 Dicranopteris linearis var. tomentosa Konahuanui summit, Southern Koolau Mountains. Growing in mostly native wet/cloud forest as a dense thicket along ridgeline along saddle between the two summit peaks. Preserved_Specimen Thomas, M.K. 316 Oahu BISH 2022-04-03
20 Dicranopteris linearis var. tomentosa 4 mi SW of Glenwood low rainforest with Metrosideros, Ilex, Scaevola, Hedyotis Preserved_Specimen Henrickson, J.S. 4066 Hawaii BISH 1969-08-06
21 Dicranopteris linearis var. tomentosa East Maui, Leeward Haleakala, Nakula NAR, southwest of Camp Release Aspect: W. Uncommon along edges of gulches. Mesic mixed koa/`ohi`a forest. Metrosideros, Acacia koa Dodonaea, Dryopteris glabra, Sadleria, Pteris, Dryopteris wallichiana, Carex, Ilex. Preserved_Specimen Pezzillo, Z. 22 Maui BISH 2022-07-19
22 Dicranopteris linearis var. tomentosa S ridge, Kipapa Gulch, Waipio, Koolau Mts dry moderate forest Preserved_Specimen Morley, H. 59 Oahu BISH 1933-12-10
23 Dicranopteris linearis var. tomentosa Munro Trail open, shrubby rainforest Preserved_Specimen Degener, O. 24276 Lanai BISH 1957-08-13
24 Dicranopteris linearis var. tomentosa 27 Miles, Glenwood scrub Preserved_Specimen Degener, O. 9180 Hawaii BISH 1929-06-29
25 Dicranopteris linearis var. tomentosa Kaala with Vaccinium and Cardamine Preserved_Specimen Takeuchi, W.N. 3070 Oahu BISH 1986-10-31
26 Dicranopteris linearis var. tomentosa Kaala with Vaccinium and Cardamine Preserved_Specimen Takeuchi, W.N. 3070 Oahu BISH 1986-10-31
27 Dicranopteris linearis var. tomentosa Puna, Volcano Road wet ohia forest Preserved_Specimen Russ, G.W. 8 Hawaii BISH 1927-07-08
28 Dicranopteris linearis var. tomentosa Kilauea with Metrosideros, Lycopodium cernuum, Cibotium, Sadleria Preserved_Specimen Degener, O. 175 Hawaii BISH 1922-12-28
29 Dicranopteris linearis var. tomentosa Lanai Shrubby rain forest Preserved_Specimen Degener, O. 26864 Lanai BISH 1960-01-18
30 Dicranopteris linearis var. tomentosa Kipapa-Waiawa Ridge, Waipio Gleichenia scrub Preserved_Specimen Grant, M.L. 7187 Oahu BISH 1934-08-07
31 Dicranopteris linearis var. tomentosa above Hiilawe woods Preserved_Specimen Rock, J.F.C. 4584 Hawaii BISH 1909-07-18
32 Dicranopteris linearis var. tomentosa Hawaii County. Puu Makaala Natural Area Reserve Site, NEON Plot PUUM_051; WGS84 MEPO5 canopy with CHTR2/ILAN mid canopy and STTA/VACA8 understory. Young lava substrate and pockets of former pasture. With Metrosideros polymorpha, Vaccinium calycinum, Coprosma sp., Cheirodendron trigynum, Styphelia tameiameiae, Anthoxanthum odoratum, Morelotia gahniiformis. Preserved_Specimen Espaniola, J. 5 Hawaii BISH 2019-09-03
33 Dicranopteris linearis var. tomentosa Puu Kaua, Southern Waianae Mountains Remnant mesic cloud forest dominated by Metrosdieros, Cheirodendron, Myrsine, Cibotium, and Diplopterygium. Preserved_Specimen Thomas, M.K. 252 Oahu BISH 2022-02-02
34 Dicranopteris linearis var. tomentosa Kilauea Caldera, outer edge open Metrosideros forest Preserved_Specimen Fosberg, F.R. 41999 Hawaii BISH 1961-08-10
35 Dicranopteris linearis var. tomentosa Manana Gulch forest clearing Preserved_Specimen Degener, O. 10428 Oahu BISH 1932-10-02
36 Dicranopteris linearis var. tomentosa Kahinahina Ridge open cloud belt Preserved_Specimen Degener, O. 30128 Lanai BISH 1963-09-09
37 Dicranopteris linearis var. tomentosa Puu Kaala, Waianae Mts dry slope Preserved_Specimen Lum, M. Oahu BISH 1933-01-08
38 Dicranopteris linearis var. tomentosa Kilauea, Byron Ledge Trail entrance to damp part Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-16
39 Dicranopteris linearis var. tomentosa West Maui, SW face of Mauna Eke with Diplopterygium pinnatum, Metrosideros polymorpha, Ilex anomala, Machaerina angustifolia, Styphelia tameiameiae, Vaccinium dentatum, Sadleria pallida Preserved_Specimen Oppenheimer, H.L. 148 Maui BISH 1996-09-28
40 Dicranopteris linearis var. tomentosa Kapoho near tree molds Preserved_Specimen Lyon, H.L. Hawaii BISH 1931-11-01
41 Dicranopteris linearis var. tomentosa Hilo, Ainako. Under Eucalyptus. Preserved_Specimen Lohman, M.L. Hawaii BISH 1948-07-20
42 Dicranopteris linearis var. tomentosa S of jct. rtes 11 and 148 lava rock, with Metrosideros and Cibotium Preserved_Specimen Stuessy, T.F. 6106 Hawaii BISH 1982-05-29
43 Dicranopteris linearis var. tomentosa Near Kaala summit on crest of ridge, upper slopes of Kamaileanu Ridge, Waianae Kai FR, Waianae Mts In wet forest of Metrosideros, Cheirodendron, Pelea etc Preserved_Specimen Nagata, K.M. 2385 Oahu BISH 1981-12-02
44 Dicranopteris linearis var. tomentosa Makua Valley, summit ridge at head dry, exposed Preserved_Specimen Degener, O. 9176 Oahu BISH 1932-06-26
45 Dicranopteris linearis var. tomentosa SE slope of Kaala open woods Preserved_Specimen Degener, O. 11541 Oahu BISH 1937-12-19
46 Dicranopteris linearis var. tomentosa Kaala, Waianae Mts bog Preserved_Specimen Hume, E.P. 551 Oahu BISH 1931-04-24
47 Dicranopteris linearis var. tomentosa East rim of Kilauea Iki Crater wet fern forest Preserved_Specimen Webster, G.L. 1698 Hawaii BISH 1948-06-15
48 Dicranopteris linearis var. tomentosa Pahoa, Puna open grassy Metrosideros forest Preserved_Specimen Degener, O. 31277 Hawaii BISH 1967-07-07
49 Dicranopteris linearis var. tomentosa above Hiilawe woods Preserved_Specimen Rock, J.F.C. 4587 Hawaii BISH 1909-07-18
50 Dicranopteris linearis var. tomentosa East Maui, Hana Forest Reserve, W of Town pasture, with Psidium, Ardisia, Metrosideros; along forest margins Preserved_Specimen Flynn, T.W. 2579 Maui BISH 1987-10-28
51 Dicranopteris linearis var. tomentosa Palawai, Lanaihale dense undergrowth in moist woods Preserved_Specimen St.John, H. 18847 Lanai BISH 1938-04-14
52 Dicranopteris linearis var. tomentosa Kipapa Gulch moist forest Preserved_Specimen Bryan Jr., E.H. 777 Oahu BISH 1934-09-18
53 Dicranopteris linearis var. tomentosa Puu Kaua. Southern Waianae mountains. Remnant mesic cloud forest dominated by Metrosideros, Cheirodendron, Myrsine, Cibotium, and others. Also growing amongst weeds such as Morella faya, Schinus terebinthifolius, Rubus rosifolius, etc. Preserved_Specimen Thomas, M.K. 252 Oahu BISH 2022-02-02
54 Dicranopteris linearis var. tomentosa S ridge of Kipapa Gulch, Koolau Mts wet Preserved_Specimen Hosaka, E.Y. 1118 Oahu BISH 1933-07-08
55 Dicranopteris linearis var. tomentosa NW corner of Honounau FR moist Cibotium forest Preserved_Specimen St.John, H. 22539 Hawaii BISH 1947-01-04
56 Dicranopteris linearis var. tomentosa Puu Kaua, east ridge, Honouliuli, Waianae Mts At summit, cloud zone. Preserved_Specimen Higgins, N.H. 50 Oahu BISH 1955-12-11
57 Dicranopteris linearis var. tomentosa Kaala, summit swamp Preserved_Specimen Takeuchi, W.N. 2347 Oahu BISH 1985-07-21
58 Dicranopteris linearis var. tomentosa Kalauao summit, Central Koolau mountains. Growing in semi boggy area on exposed ridge dominated by Leucobryum pachyphyllum, Campylopus hawaiicus, Dicranodontium porodictyon, Dichanthelium koolauense, Metrosideros polymorpha, Sadleria, Vaccinium, and others. Preserved_Specimen Thomas, M.K. 374 Oahu BISH 2022-03-19
59 Dicranopteris linearis var. tomentosa Kaala, Waianaeuka, Waianae Mts wooded ridge Preserved_Specimen St.John, H. 13387 Oahu BISH 1933-11-05
60 Dicranopteris linearis var. tomentosa Olaa forest Preserved_Specimen Skottsberg, C.J.F. 418 Hawaii BISH 1922-09-08
61 Dicranopteris linearis var. tomentosa Wright Road, Puna pasture edge, creeping in Preserved_Specimen Degener, O. 31377 Hawaii BISH 1967-01-19
62 Dicranopteris linearis var. tomentosa W Maui, Lahaina District, Lihau Ohia Mixed Mesic Shrubland in a mesic shrubland with Metrosideros, Dicranopteris, Sadleria, Lycopodium, Vaccinium reticulatum, Styphelia, Machaerina, Sphenomeris, Dodonaea, Wikstroemia, Pteridium and Gahnia Preserved_Specimen Welton, P. 1229 Maui BISH 1991-08-25
63 Dicranopteris linearis var. tomentosa Hanaula betw the reservoir and the Hanaula Springs In dense wet forest of Metrosideros, Cheirodendron, Sadleria, Lycopodium. Common in sunny places Preserved_Specimen Nagata, K.M. 1928 Maui BISH 1979-06-06
64 Dicranopteris linearis var. tomentosa Puu Kalena, Waianae Mts., Waianaeuka wooded summit Preserved_Specimen St.John, H. 22569 Oahu BISH 1947-02-16
65 Dicranopteris linearis var. tomentosa East Maui, Leeward Haleakala, Nakula NAR, southwest of Camp Release Aspect: W. Uncommon along edges of gulches. Mesic mixed koa/`ohi`a forest. Metrosideros, Acacia koa Dodonaea, Dryopteris glabra, Sadleria, Pteris, Dryopteris wallichiana, Carex, Ilex. Preserved_Specimen Pezzillo, Z. 21 Maui BISH 2022-07-19
66 Dicranopteris linearis var. tomentosa upper Kaimu Homesteads, Puna forest with Dicranopteris, Cibotium, Metrosideros, Sadleria, Straussia Preserved_Specimen Degener, O. 31508 Hawaii BISH 1968-02-12
67 Dicranopteris linearis var. tomentosa East Maui, Leeward Haleakala, Nakula NAR, southwest of Camp Release Aspect: W. Uncommon along edges of gulches. Mesic mixed koa/`ohi`a forest. Metrosideros, Acacia koa Dodonaea, Dryopteris glabra, Sadleria, Pteris, Dryopteris wallichiana, Carex, Ilex. Preserved_Specimen Pezzillo, Z. 23 Maui BISH 2022-07-19
68 Dicranopteris linearis var. tomentosa down Chain of Craters Road ~ 4.1 mi toward Kalapana from intersection with Crater Rim Rd; up side road ~2.1 mi., ~10 m NE of road Metrosideros, Sadleria, Dicranopteris forest Preserved_Specimen Newell, C.L. 242 Hawaii BISH 1966-08-02
69 Dicranopteris linearis var. tomentosa National Park, between Devil's Throat and Hiiaka Crater scrubby ohia forest Preserved_Specimen Webster, G.L. 1814 Hawaii BISH 1948-06-19
70 Dicranopteris linearis var. tomentosa Mt. Kalena Preserved_Specimen Topping, D.L. 2867 Oahu BISH 1924-09-14
71 Dicranopteris linearis var. tomentosa Olowalu, head, lateral ridge near center Preserved_Specimen Forbes, C.N. 2434 Maui BISH 1920-05-21
72 Dicranopteris linearis var. tomentosa Dupont Trail, Puu Kaala, Waianae Mts Preserved_Specimen Krajina, V.J. 620225333 Oahu BISH 1962-02-25
73 Dicranopteris linearis var. tomentosa Kaala, summit Preserved_Specimen Lamoureux, C.H. 2424 Oahu BISH 1963-10-20
74 Dicranopteris linearis var. tomentosa Kilauea, NE Ledge Preserved_Specimen Krajina, V.J. 611225052 Hawaii BISH 1961-12-25
75 Dicranopteris linearis var. tomentosa Mt. Kaala, Waianaeuka, Waianae range Preserved_Specimen Wilbur, R.L. 692 Oahu BISH 1948-04-18
76 Dicranopteris linearis var. tomentosa kipuka in 1855 flow between Olaa Flume and Halelouolu Preserved_Specimen Forbes, C.N. 677 Hawaii BISH 1915-06-03
77 Dicranopteris linearis var. tomentosa 26 Mi Preserved_Specimen Morley, H. 122 Hawaii BISH 1934-08-13
78 Dicranopteris linearis var. tomentosa Puu Alii Preserved_Specimen Munro, G.C. 215 Lanai BISH 1927-09-10
79 Dicranopteris linearis var. tomentosa East Maui, Manawainui Planeze, below Ohia camp and fence in gulch. Preserved_Specimen Welton, P. 2173008 Maui BISH 2003-08-28
80 Dicranopteris linearis var. tomentosa Puu Kaua, Waianae Mts., summit Preserved_Specimen St.John, H. 20010 Oahu BISH 1939-03-05
81 Dicranopteris linearis var. tomentosa East Maui, W side of Kaliae ridge Preserved_Specimen Hobdy, R.W. 982 Maui BISH 1981-01-21
82 Dicranopteris linearis var. tomentosa East Maui; Hwy. 36 ; opposite Waipio Bay 20.9N 156.2W Preserved_Specimen Krajina, V.J. 620420015 Maui BISH 1962-04-20
83 Dicranopteris linearis var. tomentosa W Maui Preserved_Specimen Mann, H. 495 Maui BISH 1864-01-01
84 Dicranopteris linearis var. tomentosa Mountains, E end Preserved_Specimen Forbes, C.N. 212 Lanai BISH 1913-06-01
85 Dicranopteris linearis var. tomentosa Kilauea, NE ledge Preserved_Specimen Krajina, V.J. 611225017 Hawaii BISH 1961-12-25
86 Dicranopteris linearis var. tomentosa Kohala. Preserved_Specimen Rock, J.F.C. 8420 Hawaii BISH 1910-06-01
87 Dicranopteris linearis var. tomentosa Kamaileunu ridge line above Waianae Kai Preserved_Specimen Takeuchi, W.N. 609 Oahu BISH 1983-10-02
88 Dicranopteris linearis var. tomentosa Kapua AA flow Preserved_Specimen Adee, K.T. Hawaii BISH
89 Dicranopteris linearis var. tomentosa Waianaeuka, Mt. Kaala. Preserved_Specimen Lohman, M.L. Oahu BISH 1949-05-22
90 Dicranopteris linearis var. tomentosa Makaha Valley, Waianae range Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
91 Dicranopteris linearis var. tomentosa Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
92 Dicranopteris linearis var. tomentosa Kapoho, above Kalapana Preserved_Specimen Lyon, H.L. Hawaii BISH 1931-11-01
93 Dicranopteris linearis var. tomentosa Lanai Preserved_Specimen Munro, G.C. Lanai BISH
94 Dicranopteris linearis var. tomentosa Kau, Moaula Gulch Preserved_Specimen Neal, M.C. Hawaii BISH 1927-06-01
95 Dicranopteris linearis var. tomentosa Hawaii Preserved_Specimen Seale, A. Hawaii BISH 1901-01-01
96 Dicranopteris linearis var. tomentosa Honolulu, Kapalama Heights Preserved_Specimen Mitchell, D.D. Oahu BISH 1931-11-01
97 Dicranopteris linearis var. tomentosa N. Kona, Holualoa FR, S. slope of Hualalai Preserved_Specimen Kondo, Y. Hawaii BISH 1946-07-13
98 Dicranopteris linearis var. tomentosa Puu Ohu Reservoir, Kohala Mts Preserved_Specimen Corn, C. Hawaii BISH 1984-03-27
99 Dicranopteris linearis var. tomentosa unknown Preserved_Specimen (Not on sheet) 22 BISH
100 Dicranopteris linearis var. tomentosa E slopes of Kaala Preserved_Specimen Forbes, C.N. 1612 Oahu BISH 1910-11-11
101 Dicranopteris linearis var. tomentosa L side of Chain of Craters road, just past Hilina Pali road turnoff, 7 m in Preserved_Specimen Herat, T. 1051 Hawaii BISH 1974-07-15
102 Dicranopteris linearis var. tomentosa Hookio Preserved_Specimen Munro, G.C. 149 Lanai BISH 1913-10-06
103 Dicranopteris linearis var. tomentosa unknown Preserved_Specimen Rock, J.F.C. BISH
104 Dicranopteris linearis var. tomentosa upper Makaha Preserved_Specimen Takeuchi, W.N. 2128 Oahu BISH 1985-03-03
105 Dicranopteris linearis var. tomentosa Mt. Kaala, Waianaeuka, Waianae range Preserved_Specimen Webster, G.L. 1528 Oahu BISH 1948-04-18
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